Well, without getting too detailed, the general principles are:
Ink flow is determined by nib to feed spacing, nib tine spacing, and in the case of the 51, nib to hood spacing. Of course, blocked feed channels air return channels and air leaks also play a part, but for pens in good condition, the first items I mentioned are the determining factors.
Ink is delivered to the tip of the nib by capillary action, so if the spacing is too wide, it can't 'wick' to where its needed.
After the spacing is closed to the critical point where the ink flows freely (usually the wettest flow), further narrowing of the space restricts the ink flow and it becomes a dryer writer.
In the 51, the hood/nib area is like the feed/nib area of other pens - it is used to deliver the ink to the nib by capilliary action, hence it's spacing is important - as is the tine spacing.
Writing upside down with the pen tends to push the tines closer together, and increase the hood to tine spacing. If the flow improves, this is what you want to do as an adjustment. If the flow improves when you press harder on the nib in the normal position, you'll want to spread the tines a little, or decrease the hood/nib spacing to improve flow.
Of course, experience is *very* useful in determining the adjustments to make, how much adjusting to do, and potentially how much damage is done if carried out indiscriminately...
Does that help?
Regards,
Gerry